[1] Univ Missouri, Dept Vet Pathobiol, Columbia, MO USA
[2] Univ Missouri, Dept Vet Med & Surg, Columbia, MO 65211 USA
[3] Univ Missouri, Bond Life Sci Ctr, Columbia, MO USA
[4] South Dakota State Univ, Dept Vet & Biomed Sci, Brookings, SD USA
[5] Univ Montreal, Dept Pathol & Microbiol, Montreal, PQ, Canada