A GENERIC REVISION OF THE PHALLODRILINAE (OLIGOCHAETA, TUBIFICIDAE)

The generic classification of the subfamily Phallodrilinae (with a total of 252 species) is revised according to phylogenetic principles. Morphological character patterns are assessed under the principle of maximum parsimony, but due to the poor resolution and instability in the resulting cladograms, the analysis is supplemented with intuitive weighting of synapomorphic similarities. Thirty-one genera are recognized. The former definition of the type genus Phallodrilus Pierantoni, 1902, is revised and the members of this genus are reclassified into 20 taxa, each hypothesized to be monophyletic: Phallodrilus s.str. (revised to contain the type species only). Somalidrilus gen.n., Milliganius gen.n, Phallodriloides gen.n., Albanidrilus gen.n., Abyssidrilus gen.n, Paraktedrilus gen.n, Uniporodrilus Erseus, 1979, Inermidrilus gen.n., Gianius gen.n., Mexidrilus gen.n., Peosidrilus Baker & Erseus, 1979, Atlantidrilus Erseus, 1983, Thalassodrilus Brinkhurst, 1963, Pirodrilus gen.n., Pirodriloides gen.n., Marionidrilus gen.n, Pectinodrilus gen.n., Pacifidrilus gen.n. and Pseudospiridion gen.n. Nine new species are described from the Northwest Atlantic: Phallodriloides pinnulatus sp.n., Abyssidrilus opulentus sp.n., A. potens sp.n., A. altoides sp.n., Mexidrilus obtusus sp.n. The genus Discordioprostatus Baker, 1982, is synonymized with Nootkadrilus Baker, 1982.