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- [7] Highly Prolific Booroola Sheep Have a Mutation in the Intracellular Kinase Domain of Bone Morphogenetic Protein IB Receptor (ALK-6) That Is Expressed in Both Oocytes and Granulosa Cells1[J] . Theresa Wilson,Xi-Yang Wu,Jennifer L. Juengel,Ian K. Ross,Joanne M. Lumsden,Eric A. Lord,Ken G. Dodds,Grant A. Walling,John C. McEwan,Anne R. O’Connell,Kenneth P. McNatty,Grant W. Montgomery.Biology of Reproduction . 2001 (4)