[1] Johns Hopkins Univ, Dept Biomed Engn, Baltimore, MD 21205 USA
[2] Johns Hopkins Univ, Inst Cell Engn, Baltimore, MD 21205 USA
[3] Swiss Fed Inst Technol, Dept Biosyst Sci & Engn, Basel, Switzerland
[4] Max Planck Inst Evolutionary Anthropol, Leipzig, Germany
[5] Max Planck Inst Mol Cell Biol & Genet, Dresden, Germany