[1] Univ York, Dept Biol, York YO10 5DD, N Yorkshire, England
[2] South African Environm Observat Network, ZA-3202 Cascades, South Africa
[3] Univ Cape Town, Dept Bot, ZA-7701 Rondebosch, South Africa
[4] Colorado State Univ, Nat Resource Ecol Lab, Ft Collins, CO 80523 USA
[5] Univ Botswana, Okavango Res Inst, Maun, Botswana
[6] Kruger Natl Pk, Sci Serv, ZA-1350 Skukuza, South Africa
[7] Grumeti Fund, Arusha, Tanzania
[8] Anglia Ruskin Univ, Anim & Environm Res Grp, Cambridge CB1 1PT, England
[9] Tanzania Wildlife Res Inst, Arusha, Tanzania
[10] Univ Witwatersrand, Sch Anim Plant & Environm Sci, ZA-2050 Johannesburg, South Africa
[11] Univ British Columbia, Beatty Biodivers Res Ctr, Vancouver, BC V6T 1Z4, Canada