Foraging patterns of male and female Double-crested Cormorants nesting in the Columbia River estuary

The nesting colony of Double-crested Cormorants, Phalacrocorax auritus (Lesson, 1831), on East Sand Island in the Columbia River estuary is currently the largest for this species on the Pacific Coast of North America. We used radiotelemetry to investigate the spatial and temporal foraging patterns of nesting cormorants to better understand how this colony of piscivores meets its resource needs. We determined that nesting adults tended to forage >5 km from the colony and foraging distribution was distinctly different between the sexes. On average, males commuted nearly twice the distance to forage compared with females. Females typically foraged in the estuarine mixing zone, reportedly the region of the estuary with the greatest densities of schooling fishes, while males tended to commute more than 15 km to forage in the freshwater zone. Foraging intensity of both sexes varied by time of day, tide stage, and tide series; foraging generally intensified during ebb tides. These gender differences in foraging patterns, combined with the ability to forage at considerable distance from the colony on a wide variety of prey, may allow this large and growing colony to remain productive while potentially competing for food with many thousands of other piscivorous waterbirds that use East Sand Island.