[1] Univ Cape Town, Nansen Tutu Ctr Marine Environm Res, ZA-7701 Cape Town, South Africa
[2] Univ Cape Town, MA RE Inst, Dept Oceanog, ZA-7701 Cape Town, South Africa
[3] UBO, CNRS, IRD, IFREMER,UMR 6523,LMI ICEMASA,Lab Phys Oceans, F-29280 Plouzane, France