[1] Univ Tokyo, Inst Mol & Cellular Biol, Bunkyo Ku, Tokyo 1130032, Japan
[2] Wakayama Med Sch, Dept Anat & Neurobiol, Wakayama 6408155, Japan
[3] Kansai Coll Oriental Med, Dept Anat, Kumatori, Osaka 5900433, Japan
[4] Univ Tokyo, Inst Med Sci, Minato Ku, Tokyo 1188639, Japan
[5] Iwate Univ, Fac Agr, Morioka, Iwate 0208550, Japan
[6] Teikyo Univ, Biotechnol Res Ctr, KAST, Miyamae Ku, Kawasaki, Kanagawa 2160001, Japan