[1] Univ Leeds, Sch Geog, Dept Ecol & Global Change, Leeds LS2 9JT, W Yorkshire, England
[2] Univ Utrecht, Inst Environm Biol, Dept Plant Ecol & Biodivers, NL-3508 TB Utrecht, Netherlands
[3] Univ Utrecht, Inst Environm Biol, Dept Plant Ecophysiol, NL-3508 TB Utrecht, Netherlands
[4] German Res Ctr Geosci, Helmholtz Ctr Potsdam, Dept Climate Dynam & Landscape Evolut, D-14473 Potsdam, Germany
[5] Inst Rech Dev, Lima, Peru