[1] Univ Durham, Sch Biol & Biomed Sci, Durham DH1 3LE, England
[2] Univ Milan, Dept Life Sci, I-20133 Milan, Italy
[3] Fdn Filarete, I-20139 Milan, Italy
[4] Univ Nottingham, Sch Biosci, Plant Sci Div, Loughborough LE12 5RD, Leics, England
[5] Rothamsted Res, Harpenden AL5 2JQ, Herts, England
[6] Univ Warwick, Sch Life Sci, Coventry CV4 7ES, W Midlands, England