[1] Hirosaki Univ, Fac Agr & Life Sci, Dept Biochem & Mol Biol, Hirosaki, Aomori 0368561, Japan
[2] Hirosaki Univ, Fac Sci, Dept Biol, Hirosaki, Aomori 0368561, Japan
[3] Hirosaki Univ, RNA Res Ctr, Hirosaki, Aomori 0368561, Japan
[4] Iwate Univ, United Grad Sch Agr Sci, Morioka, Iwate 0208551, Japan
[5] Tokyo Univ Agr & Technol, Dept Int Environm Agr Sci, Fuchu, Tokyo, Japan