[1] Univ Calif Berkeley, Museum Vertebrate Zool, Lab Human Evolutionary Studies, Berkeley, CA 94720 USA
[2] Univ Calif Berkeley, Dept Integrat Biol, Berkeley, CA 94720 USA
[3] Rift Valley Res Serv, Addis Ababa, Ethiopia
[4] Kent State Univ, Div Biomed Sci, Kent, OH 44242 USA
[5] Kent State Univ, Dept Anthropol, Kent, OH 44242 USA
[6] Case Western Reserve Univ, Sch Med, Dept Anat, Cleveland, OH 44106 USA
[7] Case Western Reserve Univ, Sch Med, Cleveland Museum Nat Hist, Cleveland, OH 44106 USA
[8] Univ Tokyo, Univ Museum, Bunkyo Ku, Tokyo 1130033, Japan