[1] Washington State Univ, Dept Vet Microbiol & Pathol, Pullman, WA 99164 USA
[2] Washington State Univ, UI Ctr Reprod Biol, Pullman, WA 99164 USA
[3] Washington State Univ, WSU, Pullman, WA 99164 USA
[4] Univ Santiago de Compostela, Dept Microbiol & Parasitol, Santiago De Compostela 15706, Spain
[5] Washington State Univ, Dept Vet Microbiol & Pathol, Pullman, WA 99164 USA
[6] Royal Vet & Agr Univ, Sect Fish Dis, Dept Vet Microbiol, DK-1870 Frederiksberg C, Denmark