[1] Chinese Acad Sci, Inst Hydrobiol, State Key Lab Freshwater Ecol & Biotechnol, Wuhan 430072, Peoples R China
[2] Chinese Acad Sci, Inst Hydrobiol, Key Lab Biodivers & Conservat Aquat Organisms, Wuhan 430072, Peoples R China
[3] Uppsala Univ, Dept Anim Ecol, Evolutionary Biol Ctr, SE-75272 Uppsala, Sweden