[1] Kyoto Univ, Fac Med, Dept Cell Biol, Kyoto 6068501, Japan
[2] Kyoto Univ, Fac Med, Sch Hlth Sci, Kyoto 6068501, Japan
[3] Japan Sci & Technol Corp, Solut Oriented Res Sci & Technol, Kyoto 6068501, Japan
[4] Knowledge Action Network Res Inst Inc, Kyoto 6008317, Japan
[5] Jikei Univ, Sch Med, Dept Mol Cell Biol, Inst DNA Med, Tokyo 105, Japan