[1] Univ Roma Tor Vergata, Dept Biol, I-00173 Rome, Italy
[2] Natl Inst Infect Dis IRCCS L Spallanzani, Rome, Italy
[3] Inst Gustave Roussy, INSERM, U848, F-94805 Villejuif, France
[4] Ctr Rech Cordeliers, Paris, France
[5] Hop Europeen Georges Pompidou, AP HP, Paris, France