[1] Princeton Univ, Program Atmospher & Ocean Sci, Princeton, NJ 08544 USA
[2] Natl Oceanog Ctr, Southampton, Hants, England
[3] Geophys Fluid Dynam Lab, Princeton, NJ USA
[4] Woods Hole Oceanog Inst, Woods Hole, MA 02543 USA
[5] Univ S Carolina, Marine Sci Program, Columbia, SC 29208 USA
[6] Univ S Carolina, Dept Biol Sci, Columbia, SC 29208 USA
[7] Lab Sci Climat & Environm, Gif Sur Yvette, France