[1] Univ Calif Los Angeles, Sch Med, Howard Hughes Med Inst, Los Angeles, CA 90095 USA
[2] Univ Calif Los Angeles, Sch Med, Dept Mol & Med Pharmacol, Los Angeles, CA 90095 USA
[3] Univ Calif Los Angeles, Sch Med, Dept Pathol & Lab Med, Los Angeles, CA 90095 USA
[4] Univ Wisconsin, Waisman Ctr, Dept Pathobiol Sci, Madison, WI 53705 USA
[5] Univ Wisconsin, Sch Vet Med, Madison, WI 53705 USA
[6] Yale Univ, Sch Med, Dept Genet, New Haven, CT 06510 USA