[1] Univ Lausanne, Dept Ecol & Evolut, CH-1015 Lausanne, Switzerland
[2] Michigan State Univ, BEACON Ctr Study Evolut Act, E Lansing, MI 48824 USA
[3] Univ Basel, Inst Zool, CH-4051 Basel, Switzerland
[4] Univ Montpellier 2, CNRS, Inst Sci Evolut, UMR 5554, F-34095 Montpellier 05, France
[5] Univ British Columbia, Dept Zool, Vancouver, BC V6T 1Z4, Canada