[1] Kyungpook Natl Univ, Dept Biol, Teachers Coll, Taegu 702701, South Korea
[2] Seoul Natl Univ, Sch Biol Sci, Seoul 151742, South Korea
[3] Wayne State Univ, Dept Biol Sci, Detroit, MI 48202 USA
[4] Univ Cologne, Abt Evolut Genet, Inst Genet, D-50931 Cologne, Germany