[1] Hong Kong Univ Sci & Technol, AMCE Program, Hong Kong, Hong Kong, Peoples R China
[2] Fisheries & Oceans Canada, Inst Ocean Sci, Sidney, BC V8L 4B2, Canada
[3] Univ Tokyo, Ocean Res Inst, Nakano Ku, Tokyo 1648639, Japan
[4] Tohoku Natl Fisheries Res Inst, Shiogama, Miyagi 9850001, Japan
[5] Frontier Res Syst Global Change, Kanazawa Ku, Yokohama, Kanagawa 2360001, Japan