[1] Univ Reading, Sch Biol Sci, Reading RG6 6BX, Berks, England
[2] Univ London Imperial Coll Sci Technol & Med, Div Biol, London SW7 2AZ, England
[3] Univ Reading, Sch Pharm, Reading RG6 6BX, Berks, England
[4] Royal Bot Gardens, Richmond TW9 3AB, Surrey, England