[1] RIKEN, Yokohama Inst, Genom Sci Ctr,Tsurumi Ku, Plant Funct Explorat Team,Plant Funct Genom Res G, Yokohama, Kanagawa 2300045, Japan
[2] Tokyo Univ Sci, Dept Biol Sci & Technol, Noda, Chiba 2788510, Japan
[3] RIKEN, Yokohama Inst, Plant Mutat Explorat Team, Plant Funct Genom Res Grp,Tsurumi Ku, Yokohama, Kanagawa 2300045, Japan
[4] RIKEN, Tsukuba Inst, Plant Mol Biol Lab, Tsukuba, Ibaraki 3050074, Japan
[5] Yokohama City Univ, Grad Sch Integrated Sci, Tsurumi Ku, Yokohama, Kanagawa 2300045, Japan