[1] Northwestern Univ, Sch Med, Dept Med, Chicago, IL 60611 USA
[2] Northwestern Univ, Sch Med, Dept Cell & Mol Biol, Chicago, IL 60611 USA
[3] Univ Penn, Sch Med, Abramson Family Canc Res Inst, Cell & Mol Biol Grad Grp, Philadelphia, PA 19104 USA
[4] Univ Penn, Sch Med, Howard Hughes Med Inst, Philadelphia, PA 19104 USA