It has been known for a decade that the plant MADS genes are important regulators of meristem and floral organ identity. The MADS family in Arabidopsis consists of more than 80 members and, until recently, the function of the majority of these genes was unknown. With the enhanced ability to generate loss-of-function mutants and over-expression lines, the function of the MADS gene family members is beginning to be elucidated. Recent progress demonstrates that MADS genes in Arabidopsis are important regulators not only of meristem and floral organ identity but also of flowering timing and cell-type specification in floral organs.