[1] Yale Univ, Yale Ctr Med Informat, Program Computat Biol & Bioinformat, New Haven, CT 06520 USA
[2] Yale Univ, Sch Med, Ctr Med Informat, New Haven, CT 06520 USA
[3] Yale Univ, Sch Med, Dept Genet, New Haven, CT 06520 USA
[4] Yale Univ, Dept Comp Sci, New Haven, CT 06520 USA
[5] Natl Univ Ireland Galway, Digital Enterprise Res Inst, Galway, Ireland
[6] Konrad Lorenz Inst Evolut & Cognit Res, Altenberg, Austria
[7] Yale Univ, Dept Mol Biophys & Biochem, New Haven, CT 06520 USA