[1] AZTI Tecnalia, Div Marine Res, Pasaia 20110, Spain
[2] Univ Hull, Inst Estuarine & Coastal Studies, Kingston Upon Hull HU6 7RX, N Humberside, England
[3] Natl Environm Res Inst, Dept Marine Ecol, DK-4000 Roskilde, Denmark
[4] Finnish Environm Inst SYKE, Ctr Marine Res, FI-00251 Helsinki, Finland
[5] Inst Environm & Sustainabil, EC Joint Res Ctr, I-21020 Ispra, Italy