[1] Washington Univ, Dept Pathol & Immunol, Sch Med, St Louis, MO 63110 USA
[2] Washington Univ, Sch Med, Dept Mol Microbiol, St Louis, MO 63110 USA
[3] Tokyo Med & Dent Univ, Dept Physiol & Cell Biol, Tokyo, Japan
[4] Japan Sci & Technol Agcy, Solut Oriented Res Sci & Technol, Kawaguchi, Japan
[5] Vanderbilt Univ, Sch Med, Dept Pediat, Nashville, TN 37212 USA