[1] UVA Sch Med, Dept Cell Biol, Charlottesville, VA 22908 USA
[2] Univ Virginia, Dept Microbiol, Charlottesville, VA 22908 USA
[3] Harvard Univ, Sch Med, Canc Biol Program, Boston, MA 02215 USA
[4] SUNY Upstate Med Univ, Dept Cell & Dev Biol, Syracuse, NY 13210 USA