[1] Washington Univ, Sch Med, Dept Pathol & Immunol, St Louis, MO 63110 USA
[2] Tohoku Univ, Dept Expt Immunol, Inst Dev Aging & Canc, Sendai, Miyagi 9808575, Japan
[3] RIKEN, Res Ctr Allergy & Immunol, Lab Lynphocyte Differentiat, Tsurumi Ku, Kanagawa 2300045, Japan
[4] Washington Univ, Dept Med, St Louis, MO 63110 USA
[5] Washington Univ, Dept Cell Biol, St Louis, MO 63110 USA