[1] Oregon State Univ, Coll Ocean & Atmospher Sci, Corvallis, OR 97331 USA
[2] Univ Alaska, Sch Fisheries & Ocean Sci, Fairbanks, AK 99775 USA
[3] Univ Georgia, Dept Marine Sci, Athens, GA 30602 USA
[4] Univ Washington, Sch Oceanog, Seattle, WA 98195 USA
[5] Univ Washington, Dept Earth & Space Sci, Seattle, WA 98195 USA