[1] IRD UR GEODES, F-93143 Bondy, France
[2] Ecole Normale Super, CNRS, UMR 7625, F-75230 Paris, France
[3] CHU Pitie Salpetriere, LENA, CNRS, UPR 640, F-75651 Paris, France
[4] Univ Quebec, Canada Res Chair Conservat No Ecosyst, Rimouski, PQ G5L 3A1, Canada
[5] IRD, Ctr REch Halieut Mediterranneen & Trop, F-34203 Sete, France
[6] Univ Oslo, Dept Biol, CEES, N-0316 Oslo, Norway
[7] CNRS, Ctr Etudes Biol Chize, F-79360 Villiers En Bois, France