[1] Univ Utrecht, Med Ctr, Dept Pathol, NL-3508 GA Utrecht, Netherlands
[2] Ctr Biomed Genet, Hubrecht Lab, Utrecht, Netherlands
[3] Acad Med Ctr, Dept Gastroenterol, Amsterdam, Netherlands
[4] Acad Med Ctr, Dept Pathol, Amsterdam, Netherlands
[5] Acad Med Ctr, Dept Genet Metab Dis, Amsterdam, Netherlands
[6] Acad Med Ctr, Dept Neurogenet, Amsterdam, Netherlands
[7] Johns Hopkins Univ, Sch Med, Dept Pathol, Baltimore, MD 21205 USA
[8] Johns Hopkins Univ, Sch Med, Dept Med, Div Gastroenterol, Baltimore, MD 21205 USA