[1] Univ Cape Town, Dept Oceanog, Cape Town, South Africa
[2] CSIR, Coastal Syst Res Grp, Nat Resources & Environm, Stellenbosch, South Africa
[3] Univ Cape Town, Dept Oceanog, Nansen Tutu Ctr Marine Environm Res, Cape Town, South Africa
[4] Nansen Environm & Remote Sensing Ctr, Bergen, Norway
[5] Univ Cape Town, Marine Res Inst, Dept Oceanog, Cape Town, South Africa
[6] Nelson Mandela Metropolitan Univ, Ocean Sci & Marine Food Secur, Port Elizabeth, South Africa
[7] Natl Oceanog Ctr, European Way, Southampton, Hants, England
[8] CSIR, Earth Observat Res Grp, Nat Resources & Environm, Cape Town, South Africa