[1] Kyushu Univ, Grad Sch Med Sci, Dept Pathol, Div Pathophysiol & Expt Pathol,Higashi Ku, Fukuoka 8128582, Japan
[2] Kyushu Univ, Grad Sch Med Sci, Dept Surg & Sci, Higashi Ku, Fukuoka 8128582, Japan
[3] Kyushu Univ, Grad Sch Med Sci, Dept Cardiovasc Med, Higashi Ku, Fukuoka 8128582, Japan
[4] DNAVEC Res Inc, Tsukuba, Ibaraki, Japan
[5] Natl Inst Infect Dis, Dept Viral Infect & Vaccine Control, Tokyo, Japan
[6] Natl Inst Infect Dis, AIDS Res Ctr, Tokyo, Japan