[1] Natl Inst Agrobiol Sci, Dept Plant Physiol, Tsukuba, Ibaraki 3058602, Japan
[2] Natl Inst Agrobiol Sci, Dept Mol Genet, Tsukuba, Ibaraki 3058602, Japan
[3] Univ Tokyo, Div Agr & Life Sci, Tokyo 1138657, Japan
[4] Univ Tokyo, Inst Mol & Cellular Biosci, Tokyo 1138657, Japan
[5] Natl Agr Res Ctr, Dept Rice Res, Niigata 9430193, Japan