[1] Univ Queensland, Ctr Marine Studies, St Lucia, Qld 4072, Australia
[2] Univ Queensland, Dept Earth Sci, St Lucia, Qld 4072, Australia
[3] Univ Queensland, Dept Math, St Lucia, Qld 4072, Australia
[4] Univ Queensland, Sch Life Sci, St Lucia, Qld 4072, Australia
[5] Univ Calif San Diego, Scripps Inst Oceanog, Ctr Ctr Marine Biodivers & Conservat, La Jolla, CA 92093 USA
[6] Smithsonian Trop Res Inst, Balboa, Panama
[7] Natl ctr Ecol Anal & Synth, Santa Barbara, CA USA
[8] Australian Natl Univ, Ctr Resource & Environm Studies, Canberra, ACT 0200, Australia
[9] James Cook Univ N Queensland, Sch Marine Biol, Ctr Coral Reef Biodivers, Townsville, Qld 4811, Australia
[10] Stanford Univ, Hopkins Marine Stn, Pacific Grove, CA 93950 USA
[11] Florida Inst Oceanog, St Petersburg, FL 33701 USA