[1] Univ Cape Town, Inst Infect Dis & Mol Med, Clin Sci Lab, Div Immunol, ZA-7925 Cape Town, South Africa
[2] Radboud Univ Nijmegen, Dept Internal Med, Nijmegen, Netherlands
[3] Radboud Univ Nijmegen, Nijmegen Univ Ctr Infect Dis, Nijmegen, Netherlands
[4] Univ Oxford, Sir William Dunn Sch Pathol, Oxford OX1 3RE, England
[5] Nijmegen Ctr Mol Life Sci, Tumor Immunol Lab, Nijmegen, Netherlands
[6] Natl Inst Med Res, London NW7 1AA, England
[7] Univ Aberdeen, Inst Med Sci, Sch Med Sci, Aberdeen, Scotland
[8] James H Quillen Coll Med, Dept Surg, Johnson City, TN USA