[1] Kyushu Univ, Grad Sch Med Sci, Dept Ophthalmol, Higashi Ku, Fukuoka 8128582, Japan
[2] Kyushu Univ, Grad Sch Med Sci, Dept Med Biochem, Fukuoka 8128582, Japan
[3] Kyushu Univ, Grad Sch Med Sci, Collabo Stn 2, Fukuoka 8128582, Japan
[4] Kurume Univ, Res Ctr Innovat Canc Therapy Century 21, COE, Program Med Sci, Fukuoka, Japan
[5] Harvard Univ, Sch Med, Schepens Eye Res Inst, Boston, MA USA
[6] Harvard Univ, Sch Med, Dept Ophthalmol, Massachusetts Eye & Ear Infirm, Boston, MA USA