[1] Rothamsted Res, Plant & Invertebrate Ecol Dept, Harpenden AL5 2JQ, Herts, England
[2] Univ New S Wales, Australian Def Force Acad, Sch Phys Environm & Math Sci, Canberra, ACT 2600, Australia
[3] Univ Canberra, Inst Appl Ecol, Canberra, ACT 2601, Australia
[4] Univ Greenwich, Nat Resources Inst, Chatham ME4 4TB, Kent, England