[1] Univ Nottingham, Sch Biomed Sci, Nottingham NG7 2UH, England
[2] Univ Nottingham Med Sch, Inst Neurosci, Nottingham NG7 2UH, England
[3] Univ Kent & Greenwich Medway, Medway Sch Pharm, Kent ME4 4TB, England
[4] Univ Dundee, Ninewells Hosp and Med Sch, Inst Neurosci, Div Pathol & Neurosci, Dundee, Scotland