[1] Tohoku Univ, Grad Sch Agr Sci, Dept Life Sci, Aoba Ku, Sendai, Miyagi 9818555, Japan
[2] Yale Univ, Dept Mol Cellular & Dev Biol, OML 451, New Haven, CT 06520 USA
[3] Kansas State Univ, Div Biol, Mol Cellular & Dev Biol Program, Manhattan, KS 66506 USA