[1] Univ Queensland, Australian Ctr Int & Trop Hlth & Nutr, Mol Parasitol Unit, Brisbane, Qld 4029, Australia
[2] Queensland Inst Med Res, Brisbane, Qld 4006, Australia
[3] James Cook Univ N Queensland, Townsville, Qld 4811, Australia
[4] Kochi Med Sch, Environm Hlth Sci Lab, Nankoku, Kochi, Japan
[5] Nat Hist Museum, Dept Zool, London SW7 5BD, England
[6] USDA, Inst Livestock & Poultry Sci, Beltsville, MD 20705 USA