[1] INRA, Unite Rech Pluridisciplinaire Prairies & Plantes, UR4, F-86600 Lusignan, France
[2] INRA, Unite Mixte Rech Genet Vegetale, Gif Sur Yvette, France
[3] Inner Mongolia Agr Univ, Coll Agron, Hohhot, Inner Mongolia, Peoples R China
[4] Aberystwyth Univ, Inst Biol Environm & Rural Sci, Aberystwyth, Ceredigion, Wales