[1] Chinese Acad Sci, Inst Tibetan Plateau Res, Key Lab Continental Collis & Plateau Uplift, Beijing 100101, Peoples R China
[2] Chinese Acad Sci, Ctr Excellence Tibetan Plateau Earth Sci, Beijing 100101, Peoples R China
[3] Chinese Acad Sci, Inst Bot, State Key Lab Systemat & Evolutionary Bot, Beijing 100093, Peoples R China
[4] Open Univ, Sch Environm Earth & Ecosyst Sci, Milton Keynes MK7 6AA, Bucks, England
[5] Birbal Sahni Inst Paleobot, Lucknow 226007, Uttar Pradesh, India
[6] Univ Calcutta, Dept Bot, Ctr Adv Study, Kolkata 700019, India
[7] Sidho Kanho Birsha Univ, Dept Bot, Ranchi Rd, Purulia 723104, India