[1] Seoul Natl Univ, Sch Agr Biotechnol, Suwon 441744, South Korea
[2] Korea Ginseng & Tobacco Res Inst, Taejon 305345, South Korea
[3] Korea Res Inst Biosci & Biotechnol, Taejon 305600, South Korea
[4] Chungnam Natl Univ, Dept Agr Biol, Taejon 305764, South Korea