[1] Whitehead Inst Biomed Res, Cambridge, MA 02142 USA
[2] Novartis Inst BioMed Res, Basel, Switzerland
[3] Massachusetts Gen Hosp, Brigham & Womens Hosp, Enterprise Res IS & Informat, Boston, MA 02114 USA
[4] Vanderbilt Univ, Ctr Human Genet Res, Nashville, TN USA