[1] Univ Western Australia, Western Australian Inst Med Res, Lab Mol Neurosci, Perth, WA 6000, Australia
[2] Univ Western Australia, Med Res Ctr, Perth, WA 6000, Australia
[3] Monash Med Ctr, Inst Reprod & Dev, Clayton, Vic 3168, Australia
[4] Millennium Pharmaceut Inc, Cambridge, MA USA