[1] Univ Alabama Birmingham, Sch Optometry, Vis Sci Res Ctr, Dept Physiol Opt, Birmingham, AL 35294 USA
[2] Washington Univ, Sch Med, Dept Ophthalmol & Visual Sci, St Louis, MO 63110 USA
[3] Univ Michigan, Kellogg Eye Ctr, Dept Ophthalmol, Ann Arbor, MI 48105 USA
[4] Univ Michigan, Kellogg Eye Ctr, Dept Human Genet, Ann Arbor, MI 48105 USA
[5] Johns Hopkins Med Sch, Dept Ophthalmol, Baltimore, MD 21287 USA
[6] Johns Hopkins Med Sch, Dept Mol Biol & Genet, Baltimore, MD 21287 USA
[7] Johns Hopkins Med Sch, Dept Neurosci, Baltimore, MD 21287 USA