Paleogene events in Central Eurasia: their role in the flora and vegetation cover evolution, migration of phytochore boundaries, and climate changes

The flora and vegetation of Central Eurasia evolved in the Paleogene to a significant extent in line with the scenario similar to the Late Cretaceous one. The position of high-rank phytochores was controlled by the global climatic zonality, while development stages of the flora depended on interaction between the Arctic and Tethyan water masses and direction of atmospheric flows and were determined by principal geological and paleogeographic events in the Paleogene history of Central Eurasia. Five main stages are definable in development of the Paleogene flora: (1) early-middle Danian with the wide distribution of temperate-thermophilic floras in the middle and high latitudes and their westward and southward expansion from the Pacific and Arctic regions of the Boreal realm; (2) Late Paleocene-Early Eocene with the maximal advancement of the Tethyan flora to the high latitudes and northward migration of phytochore boundaries in response to intense water exchange between the Tethys and Atlantic oceans with its trade currents and atmospheric heat transfer directly from the tropical zone in absence of the Alpine-Himalayan orogen; (3) Lutetian with development of subtropical monsoon-type floras under influence of the water mass exchange between the Arctic Basin and Peritethys with the monsoon-induced currents and atmospheric heat transfer from the Peritethys under conditions of the restricted connection between the Central Asia basins and Tethys; (4) (?) late Lutetian-Priabonian reflecting the climate inversion due to isolation of the West Siberian Sea from the Arctic Basin against the background of its continuing connection with the Peritethys; the formation of the semiclosed West Siberian Sea at that time was accompanied by development of a climate with humid winters, hot dry summers, and deficiency of average annual precipitation in the middle latitudes of Central Eurasia, where luxuriant subtropical Quercus-Laurus forests with Castanopsis that prevailed at the preceding stage were replaced by sclerophyllous arboreal-frutescent maquis; (5) Oligocene marked by the formation of the temperate deciduous mesophyllous coniferous-broad-leaved Turgai flora after definitive desiccation of the West Siberian Sea and Turgai Seaway due to global regression induced by glaciation (transition from the "warm" to "cold" biosphere).